Fish physiology, particularly nutritional biochemistry of larval stages of freshwater species cyprinids, coregonines, salmonids, esocids, and percids
Fish nutrition, particularly metabolism and requirement of ascorbic acid and tocopherol.
Evolutionary aspects of ascorbic acid synthesis in aquatic animals' aquaculture methods in reproduction and rearing of freshwater fishes with emphasis on fish bioenergetics in intensive systems, ponds, and lake cages
Sperm physiology, mechanisms of sperm-egg interaction in lamprey, sturgeon and teleost fish.
